Parting the waters. Middle Palaeolithic archaeology in the central Ionian Sea


  • Nena Galanidou



This paper sets out a conceptual framework based on the idea of connectivity, and the research design that informs a series of surveys and excavations in the central Ionian Sea targeting the Palaeolithic record. It highlights the importance of mapping the now submerged topography to get a better understanding of the relevant palaeogeography, and its wider implications for hominin settlement, landscape preferences and pathways of dispersal and expansion from mainland Greece into the Inner Ionian Archipelago and Lefkas. It argues that the sea and the offshore islands are not marginal or irrelevant but central to an understanding of Palaeolithic settlement and land use, especially during periods of low sea level. At these times, many islands would have become hills in an extensive coastal plain, representing ideal habitats for hunters and gatherers.


Baika, K. 2008. Archaeological indicators of relative sea-level changes in the Attico-Cycladic massif: preliminary results. Bulletin of the Geological Society of Greece 42(II): 33–48.

Bailey, G.N. and N.C. Flemming 2008. Archaeology of the continental shelf: marine resources, submerged landscapes and underwater archaeology. Quaternary Science Reviews 3(24): 2153–65.

Bailey, G.N., D. Sakellariou and members of the SPLASHCOS network 2012. SPLASHCOS: Submerged Prehistoric Archaeology and Landscapes of the Continental Shelf. Antiquity Project Gallery 86 (334) December.

Bailey G.N., N. Galanidou, H. Jöns, F. Lüth and H. Peeters in press. The Prehistoric Archaeology of Europe’s Submerged Landscapes: An Introduction. In Bailey G.N., N. Galanidou, H. Jöns, F. Lüth and H. Peeters (eds) The Prehistoric Archaeology of Europe’s Submerged Landscapes (Coastal Research Library). Heidelberg: Springer.

Berger, L.R., J. Hawks, D.J. de Ruiter, S.E. Churchill, P. Schmid, L.K. Delezene, T.L. Kivell, H.M. Garvin, S.A. Williams and J.M. DeSilva 2015. Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa. eLife 4:e09560.

Beyin, A. 2011. Upper Pleistocene human dispersals out of Africa: A review of the current state of the debate. International Journal of Evolutionary Biology Article ID 615094.

Borgna E. and S. Müller-Celka (eds) 2011. Ancestral Landscapes. Burial Mounds in the Copper and Bronze Ages (Central and Eastern Europe - Balkans - Adriatic - Aegean, 4th-2nd millennium B.C.) (Travaux de la Maison de l’Orient et de la Méditerranée 58). Lyon: Maison de l’Orient et de la Méditerranée.

Broodbank, C. 2014. So... what? Does the paradigm currently want to budge so much? Journal of Mediterranean Archaeology 27(2): 267–72.

Bulbeck, D. 2007. Where river meets sea: a parsimonious model for Homo sapiens colonization of the Indian Ocean Rim and Sahul. Current Anthropology 48(2): 315–21.

Chatzimpaloglou, P. 2014. Cherts and Siliceous Pre-historic Artifacts from Meganisi Island, Lefkas. M.Sc. Thesis, University of Athens.

Chavallion, J., N. Chavallion and F. Hours 1967. Industries paleolithiques de l’Elide. I –Region d’Amalias. Bulletin de Correspondance Hellenique 91: 151–201.

Dennell, R.W. 2015. Homo sapiens in China 80,000 years ago. Nature 526: 696–9. doi:10.1038/nature15640.

Dennell, R.W. and W. Roebroeks 2005. An Asian perspective on early human dispersal from Africa. Nature 438: 1099–1104.

Dousougli, Α. 1999. Palaeolithic Lefkas, in G.N. Bailey, E. Adam, E. Panagopoulou, C. Perlès and K. Zachos (eds) The Palaeolithic Archaeology of Greece and Adjacent Areas: Proceedings of the ICOPAG Conference, Ioannina, September (British School at Athens Studies 3): 288–92. London: British School at Athens.

Efstratiou, N. 2001. The Neolithic settlement of Agios Petros in Kyra-Panagia, Halonnesos and the island habitation patterns in the Aegean –a reconsideration. In A. Sampson (ed.) Archaeological Research in the Northern Sporades (in Greek): 231–50. Alonnesos: Municipality of Alonnesos.

Emiliani, C. 1955. Pleistocene temperatures. Journal of Geology 63: 538–78.

Evans, J.D. 2008. Approaching the Labyrinth, in V. Isaakidou and P. Tomkins (eds) Escaping the Labyrinth: the Cretan Neolithic in Context (Sheffield Studies in Aegean Archaeology 8): 11–20. Oxford: Oxbow Books.

Evans, J.D. and C. Renfrew 1968. Excavations at Saliagos near Antiparos (The British School at Athens Supplementary Volume 5). London: Thames & Hudson.

Faure, H., R.C. Walter and D.R. Grant 2002. The coastal oasis: ice age springs on emerged continental shelves. Global and Planetary Change 33(1–2): 47–56.

Ferentinos, G., M. Gkioni, M. Geraga and G. Papatheodorou 2012. Early seafaring activity in the southern Ionian Islands, Mediterranean Sea. Journal of Archaeological Science 39(7): 2167–76.

Flemming, N.C. 1973. Archaeological evidence for eustatic and tectonic components of relative sea level change in the South Aegean, in D.J. Blackman (ed.) Marine Archaeology (Colston Symposium Papers 23): 1–66. London, Butterworth.

Flemming, N.C. 1978. Holocene eustatic changes and coastal tectonics in the northeast Mediterranean: implications for models of crustal consumption. Philosophical Transactions of the Royal Society of London 289(1362): 405–58.

Flemming, N.C. 1983. Preliminary geomorphological survey of an early Neolithic submerged site in the Sporadhes, N. Aegean, in P.M. Masters and N.C. Flemming (eds) Quaternary Coastlines and Marine Archaeoloogy: 233–68. London and New York: Academic Press.

Flemming, N., M.N. Çağatay, F.L. Chiocci, N. Galanidou, H. Jöns, G. Lericolais, T. Missiaen, F. Moore, A. Rosentau, D. Sakellariou, B. Skar, A. Stevenson and H. Weerts 2014. Land Beneath the Waves: Submerged Landscapes and Sea Level Change. A Joint Goescience-humanities Strategy for European Continental Shelf Prehistoric Research, in N.C. Chu and N. McDonough (eds) European Marine Board Position Paper 21. Ostend: European Marine Board.

Galanidou, N. 1996. ‘So far away, so close’: Historical and theoretical perspectives on Palaeolithic and Mesolithic research in Greece. Ariadne 8: 7–29.

Galanidou, N. 2014a. Advances in the Palaeolithic and Mesolithic archaeology of Greece for the new millennium. Pharos XX(I): 1–40.

Galanidou, N. 2014b. Archaic hominins on Crete: fact or fiction? Journal of Mediterranean Archaeology 27(2): 260–7.

Galanidou, N. 2014c. Archaeological survey on the Inner Ionian Sea archipelago, in C. Smith (ed.) Encyclopaedia of Global Archaeology: 3882–8. New York: Springer.

Galanidou, N. 2015. Seascape survey on the Inner Ionian Sea archipelago, in M. Carver, B. Gaydarska and S. Mondon-Subias (eds) Field Archaeology from Around the World. Ideas and Approaches: 101–6. New York: Springer.

Galanidou, N. 2016. Before ancient Nericos, what? Palaeolithic finds from Karyotes Lefkas, in C. Papadatou-Giannopoulou (ed.) Nericos, Lefkas, Kastro the Longest-lived Capital of Lefkas, Conference Proceedings August 2010 (in Greek): 83–111. Lefkas: Cultural Centre.

Galanidou, N., G. Iliopoulos and C. Papoulia 2016a. The Palaeolithic settlement of Lefkas: Archaeological evidence in a palaeogeographic context. Journal of Greek Archaeology 1: 1–32.

Galanidou, N., C. Athanassas, J. Cole, G. Iliopoulos, A. Katerinopoulos, A. Magganas and J. McNabb 2016b. The Acheulean site at Rodafnidia, Lisvori on Lesbos, Greece: part I, in K. Harvati and M. Roksandic (eds) Paleoanthropology of the Balkans and Anatolia: Human Evolution and its Context. Springer: Dordrecht.

Galanidou, N., O. Vikatou, M. Gatsi-Stavropoulou, A. Vasilakis, G. Iliopoulos, V. Staikou, M. Veikou, J. Forsén, C. Morgan, J. Vroom, C. Papoulia, P. Zervoudakis and K. Prasas 2017a. The Archaeological Survey in the Inner Ionian Sea Archipelago, in O. Vikatou (ed.) Archaeological Work in Aetoloakarnania and Lefkas from Prehistoric to Roman Times, Proceedings of the 2nd International Archaeological and Historical Conference, 6–8 December 2013, Mesolonghi (in Greek): 277–305. Mesolonghi.

Galanidou, N., G. Iliopoulos, C. Papoulia and V. Staikou 2017b. Palaeolithic Lefkas: Recent research, in O. Vikatou (ed.) Archaeological Work in Aetoloakarnania and Lefkas from Prehistoric to Roman Times, Proceedings of the 2nd International Archaeological and Historical Conference, 6–8 December 2013, Mesolonghi (in Greek): 343–359. Mesolonghi.

Galanidou N., K. Dellaporta and D. Sakellariou in press. The Prehistoric Archaeology of the Greek Continental Shelf, in Bailey G.N., N. Galanidou, H. Jöns, F. Lüth and H. Peeters (eds) The Prehistoric Archaeology of Europe’s Submerged Landscapes (Coastal Research Library). Springer.

Galanidou, N. and D. Nazou in press. Archaeology and tourism at Meganisi, central Ionian sea, in N. Galanidou (ed.) Essays in Public Archaeology (in Greek). Athens: Kaleidoscope editions.

Gamble, C. 1999. The Palaeolithic Settlement of Europe. Cambridge: Cambridge University Press.

Gamble, C. 2007. Origins and Revolutions. Human Identity in Earliest Prehistory. Cambridge: Cambridge University Press.

Gamble, C. 2013. Settling the Earth. The Archaeology of Deep Human History. Cambridge: Cambridge University Press.

Harff, J., G.N. Bailey and F. Lüth (eds) 2016. Geology and Archaeology: Submerged Landscapes of the Continental Shelf, Geological Society of London Special Publication 411/DISPERSE contribution 25.

Harmand, S. et al. 2015. 3.3-million-year-old stone tools from Lomekwi 3, West Turkana, Kenya. Nature 521: 310–15 (21 May 2015)

Hays, J.D., J. Imbrie and N.J. Shackleton 1976. Variations in the Earth’s Orbit: Pacemaker of the Ice Ages. Science 194(4270): 1121–32.

Head, M.J., P.L. Gibbard and T. van Kolfschoten 2015. The Quaternary system and its formal subdivision. Quaternary International 383:1–3.

Henderson, J., C. Gallou, N.C. Flemming and E. Spondylis 2011. The submerged prehistoric town at Pavlopetri in southern Greece, in J. Benjamin, C. Bonsall, C. Pickard and A. Fischer (eds) Underwater Archaeology and the Submerged Prehistory of Europe: 207–18. Oxford: Oxbow.

Hershkovitz, I., G.W. Weber, R. Quam, M. Duval, R. Grün, L. Kinsley, A. Ayalon, M. Bar Matthews, H. Valladas, N. Mercier, J.L. Arsuaga, M. Martinón-Torres, J.M. Bermúdez de Castro, C. Fornai, L. Martín-Francés, R. Sarig, H. May, V.A. Krenn, V. Slon, L. Rodríguez, R. García, C. Lorenzo, J.M. Carretero, A. Frumkin, R. Shahack-Gross, D. Bar-Yosef Mayer, Y. Cui, X. Wu, N. Peled, I. Groman-Yaroslavski, L. Weissbrod, R. Yeshurum, A. Tsatsakin, Y. Zaidner and M. Weinstein-Evron 2018. The earliest modern humans outside Africa. Science 359(6374): 456–59.

Hublin, J. J., A. Ben-Ncer, S.E. Bailey, S.E. Freidline, S. Neubauer, M.M. Skinner, I. Bergmann, A. Le Cabec, S. Bernazzi, K. Harvati and P. Gunz 2017. New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature 546(7657): 289–92.

Hsü, K.J. 1982. The Mediterranean was a Desert: a Voyage of the Glomar Challenger. New Jersey: Princeton University Press.

Imbrie, J. and K. Palmer Imbrie 1979. Ice Ages. Solving the Mystery. Cambridge, MA and London: Harvard University Press.

Jöris, O. 2014. Early Palaeolithic Europe, in C. Renfrew and P. Bahn (eds) The Cambridge World Prehistory, III: 1703–46. Cambridge: Cambridge University Press.

Kohn, M. 2006. Made in Savannahstan. New Scientist 191: 34–9.

Koutsoumpos, L. and N. Galanidou 2015. Catching water: traditional water-collecting and storing structures at Meganisi. Paper read at Conference BIOCULTURAL 2015: International Conference on Sustainability in Architectural Cultural Heritage.

Lambeck, K. 1996. Sea-level change and shoreline evolution in Aegean Greece since Upper Palaeolithic time. Antiquity 70: 588–611.

Lambeck, K. and A. Purcell 2005. Sea-level change in the Mediterranean Sea since the LGM: model predictions for tectonically stable areas. Quaternary Science Reviews 24: 1969–88.

Leppard, T. 2014. Modeling the impacts of Mediterranean island colonization by Archaic Hominins: the likelihood of an insular Lower Palaeolithic. Journal of Mediterranean Archaeology 27: 231–53.

Leppard, T. 2015. Passive dispersal versus strategic dispersal in island colonization by hominins. Current Anthropology 56(4): 590–5.

Leroi-Gourhan, A. 1964. Découvertes paléolithiques en Élide. Bulletin de Correspondence Hellénique 88(2): 1–8.

Lykousis, V. 2009. Sea-level changes and shelf break prograding sequences during the last 400 ka in the Aegean margins: Subsidence rates and palaeogeographic implications. Continental Shelf Research 29: 2037–44.

Magganas, A., N. Galanidou, P. Chatzimpaloglou, M. Kati, G. Iliopoulos and A. Katerinopoulos in press. Petrology and Provenance of lithic raw materials used to knap stone: a case study from the Inner Ionian Sea. The Bulletin of the Geological Society of Greece

Morrison, I.A. 1968. Appendix I. Relative sea level change in the Saliagos area since Neolithic times, in C. Renfrew and J.D. Evans (eds) Excavations at Saliagos Near Antiparos, British School at Athens Supplementary Volume 5: 92–98. London: British School at Athens.

Nigst, P.R., P. Haesearts, F. Damblon, C. Frank-Fellner, C. Mallol,B. Viola, M. Götzinger, L. Niven, G. Trnka and J.-J.Hublin 2014. Early modern human settlement of Europe north of the Alps occurred 43,500 years ago in a cold steppe-type environment. PNAS 111(40): 14394–14399.

Phoca-Cosmetatou, N. and R. J. Rabett 2014. Reflections on Pleistocene island occupation. Journal of Mediterranean Archaeology 27: 255–9.

Posth, C., C. Wißing, K. Kitagawa, L. Pagani, L. Van Holstein, F. Racimo, K. Wehrberger, N.J. Conard, C.J. Kind, H. Bocherens and J. Krause 2017. Deeply divergent archaic mitochondrial genome provides lower time boundary for African gene flow into Neanderthals. Nature Communications 8 (May) 16046: 1–9

Poulos, S.E., G. Ghionis and H. Maroukian 2009. Sea-level rise trends in the Attico-Cycladic region (Aegean Sea) during the last 5000 years. Geomorphology 107(1–2): 10–17.

Psychoyos, O. 1988. Déplacements de la ligne de rivage et sites archéologiques dans les régions côtières de la mer Égée, au Néolithique et à l’Age du Bronze (Studies in Mediterranean Archaeology and Literature Pocket-book 62). Jonsered: Paul Aström.

Rohling, E.J., K. Grant, M. Bolshaw, A.P. Roberts, M. Siddall, C. Hemleben and M. Kucera 2009. Antarctic temperature and global sea level closely coupled over the past five glacial cycles. Nature Geoscience 2: 500–4.

Runnels, C. 2014. Early Palaeolithic on the Greek Islands? Journal of Mediterranean Archaeology 27: 211–30.

Sakellariou, D. and N. Galanidou 2016. Pleistocene submerged landscapes and Palaeolithic archaeology in the tectonically active Aegean region, in J. Harff, G.N. Bailey and F. Lüth (eds) Geology and Archaeology: Submerged Landscapes of the Continental Shelf, Geological Society of London Special Publication 411/DISPERSE contribution 25: 145–78.

Sakellariou, D. and N. Galanidou 2017. Aegean Pleistocene landscapes above and below sea level: palaeogeographic reconstruction and hominin dispersals, in G.N. Bailey, J. Harff, and D. Sakellariou (eds) Under the Sea: Archaeology and Palaeolandscapes of the Continental Shelf: 335–59. Springer.

Sakellariou, D., V. Lykousis, M. Geraga, G. Rousakis and T. Soukisian 2017. Late Pleistocene environmental factors of the Aegean region (Aegean sea including the Hellenic Arc) and the identification of potential example areas for seabed prehistoric sites and landscapes, in N.C. Flemming, J. Harff, D. Moura, A. Burgess and G. Bailey (eds) Quaternary Palaeo-environments on the European Continental Shelf: Survival and Destruction of Prehistoric Remains. Wiley-Blackwell: 405–29.

Sordinas, A. 1969. Investigations of the prehistory of Corfu during 1964–1966. Balkan Studies 10: 393–424.

Sordinas, A. 1983. Quaternary shorelines in the region of Corfu and adjacent islets, western Grecce, in P.M. Masters and N.C. Flemming (eds) Quaternary Coastlines and Marine Archaeology, 335–44. London and New York: Academic Press.

Stiros, S.C., P.A. Pirazzoli and M. Fontugne 2009. New evidence of Holocene coastal uplift in the Strophades Islets (W Hellenic Arc, Greece). Marine Geology 267: 207–11.

Tzortzakaki, O. 2012. Diversity of terrestrial macrofauna on Meganisi and its satellite islets, Central Ionian Sea. M.Sc. Dissertation, University of Patras.

van Andel, T.H. and J.C. Shackleton 1982. Late Palaeolithic and Neolithic coastlines of Greece and the Aegean. Journal of Field Archaeology 9: 445–54.

van Andel, T. and C. Runnels 2005. Karstic wetland dwellers of Middle Palaeolithic Epirus. Journal of Field Archaeology 30: 367-384.

Vött, A. 2007. Relative sea level changes and regional tectonic evolution of seven coastal areas in NW Greece since the mid-Holocene. Quaternary Science Reviews 26(7–8): 894–919.

Walker, M.J.C., M. Berkelhammer, S. Björck, L.C. Cwynar, D.A. Fisher, A.J. Long, J.J. Lowe, R.M. Newnham, S.O. Rasmussen and H. Weiss 2012. Formal subdivision of the Holocene Series/Epoch: a discussion paper by a working group of INTIMATE (Integration of ice-core, marine and terrestrial records) and the Subcommission on Quaternary Stratigraphy (International Commission on Stratigraphy). Journal of Quaternary Science 27(7): 649–59.

Westley, K. and J. Dix 2006. Coastal environments and their role in prehistoric migrations. Journal of Maritime Archaeology 1(1): 9–28.

Winder, I.C., G.C.P. King, M. Devès and G.N. Bailey 2013. Complex topography and human evolution: the missing link. Antiquity 87: 333–49.

Zachos, K. and A. Dousougli 2003. Lefkas. Historical-Archaeological Review through the Exhibits of the Archaeological Museum (in Greek). Athens: Ministry of Culture, 12th EPCA.

Zavitsanou, A. 2016. Paleogeographical Reconstruction of the Northern Inner Ionian Sea Archipelago during Late Quaternary Low Sea Level Periods. M.Sc. Dissertation, University of Athens.

Zavitsanou, A., D. Sakellariou, G. Rousakis, P. Georgiou and N. Galanidou 2015. Paleogeographic reconstruction of the Inner Ionian Sea during Late Pleistocene low sealevel stands: preliminary results. 11th Panhellenic Symposium on Oceanography and Fisheries, Mytilene, Lesvos Island, Greece: 997–1000.

Zavitsanou, A., D. Sakellariou, G. Rousakis, P. Georgiou, N. Galanidou and J. Alexopoulos in prep. Paleogeographic reconstruction of the northern Inner Ionian Sea Archipelago (Western Greece) during Late Quaternary low sea level periods, paper submitted to Quaternary Science Reviews.



How to Cite

Galanidou, N. (2018). Parting the waters. Middle Palaeolithic archaeology in the central Ionian Sea. Journal of Greek Archaeology, 3, 1–23.



Prehistory and Protohistory